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Details on Person UniProt:Q8IYL9 GPR65

Class:IdReferenceGeneProduct:99191
_chainChangeLogchain:1-337 added on Sat February 7 2015
_displayNameUniProt:Q8IYL9 GPR65
_timestamp2026-02-20 21:30:24
chainchain:1-337
checksumCFDBF217291C0FA2
commentFUNCTION Proton-sensing G-protein coupled receptor activated by extracellular pH, which is required to monitor pH changes and generate adaptive reactions (PubMed:15326175, PubMed:15618224, PubMed:20855608, PubMed:33478938, PubMed:37722051, PubMed:39753132, PubMed:40211064). Activated by an optimal pH of 7.4 (PubMed:39753132). Ligand binding causes a conformation change that triggers signaling via guanine nucleotide-binding proteins (G proteins) and modulates the activity of downstream effectors, such as adenylate cyclase (PubMed:15326175, PubMed:15618224, PubMed:37722051, PubMed:39753132). GPR65 is mainly coupled to G(s) G proteins and mediates activation of adenylate cyclase activity (PubMed:15618224, PubMed:37722051, PubMed:39753132). May also act as a receptor for the glycosphingolipid psychosine (PSY) and several related glycosphingolipids (PubMed:11309421, PubMed:15326175). Plays a role in immune response by maintaining lysosome function and regulating T-cell metabolism (PubMed:27287411). Acts as a regulator of inflammation by mediating pH-sensing of extracellular acidification which takes place in inflamed tissues: activation regulates endo-lysosomal function of immune cells and T-cell metabolism (By similarity). Constitutively active in endosomes and stimulates adenylate cyclase production from endosomes independently from extracellular pH changes (PubMed:39753132).ACTIVITY REGULATION Activated by a network of residues that connects an extracellular-facing cavity to Glu-142, a conserved charged residue buried in the transmembrane core of the receptor (PubMed:39753132). Protonation likely drives conformational changes in extracellular loop 2 (ECL2), which stabilizes movement of transmembrane 3 (TM3) and a series of rearrangements that connect the extracellular-facing cavity to Glu-142, a residue only conserved in proton-sensing G-protein coupled receptors (PubMed:39753132). Activated by BTB09089, a positive allosteric modulator (PubMed:26550826).SUBCELLULAR LOCATION Internalizes and localizes to early and late endosomes, from where GPR65 signals at steady state, irrespective of extracellular pH (PubMed:39753132). Changes in extracellular pH may relocalize receptor signaling to the cell membrane (PubMed:39753132).TISSUE SPECIFICITY Predominantly expressed in thymus, spleen, lymph nodes, small intestine, lung, placenta and peripheral blood leukocytes.DOMAIN A multitude of proton-sensing residues, which include extracellular histidine (His-10, His-14 and His-243) residues or triad of buried acidic residues (Asp-60, Glu-142, Asp-286), contribute to activation of the G-protein coupled receptor activity and pH sensitivity.SIMILARITY Belongs to the G-protein coupled receptor 1 family.
descriptionrecommendedName: fullName evidence="16"G-protein coupled receptor 65 alternativeName: fullName evidence="15"Psychosine receptor alternativeName: fullName evidence="17"T-cell death-associated gene 8 protein
geneNameGPR65
TDAG8
identifierQ8IYL9
isSequenceChangedFALSE
keyword3D-structure
Apoptosis
Cell membrane
Disulfide bond
Endosome
G-protein coupled receptor
Glycoprotein
Membrane
Proteomics identification
Receptor
Reference proteome
Transducer
Transmembrane
Transmembrane helix
modified[InstanceEdit:9836292] Weiser, Joel, 2023-05-25
[InstanceEdit:9852000] Weiser, Joel, 2023-11-03
[InstanceEdit:9926675] Weiser, Joel, 2024-11-03
[InstanceEdit:9963647] Weiser, Joel, 2025-08-15
[InstanceEdit:9983091] Weiser, Joel, 2026-02-20
nameGPR65
referenceDatabase[ReferenceDatabase:2] UniProt
referenceGene[ReferenceDNASequence:8993843] ENSEMBL:ENSG00000140030 GPR65 [Homo sapiens]
secondaryIdentifierGPR65_HUMAN
O75819
sequenceLength337
species[Species:48887] Homo sapiens
(referenceEntity)[EntityWithAccessionedSequence:444624] GPR65 [plasma membrane] [Homo sapiens]
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No pathways have been reviewed or authored by UniProt:Q8IYL9 GPR65 (99191)