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Details on Person UniProt:O60563 CCNT1

Class:IdReferenceGeneProduct:69859
_chainChangeLogchain:1-726 added on Fri February 6 2015
_displayNameUniProt:O60563 CCNT1
_timestamp2024-11-03 20:18:17
chainchain:1-726
checksum4637EFB2DDEDFE13
commentFUNCTION Regulatory subunit of the cyclin-dependent kinase pair (CDK9/cyclin-T1) complex, also called positive transcription elongation factor B (P-TEFb), which facilitates the transition from abortive to productive elongation by phosphorylating the CTD (C-terminal domain) of the large subunit of RNA polymerase II (RNA Pol II) (PubMed:16109376, PubMed:16109377, PubMed:30134174, PubMed:35393539). Required to activate the protein kinase activity of CDK9: acts by mediating formation of liquid-liquid phase separation (LLPS) that enhances binding of P-TEFb to the CTD of RNA Pol II (PubMed:29849146, PubMed:35393539).FUNCTION (Microbial infection) In case of HIV or SIV infections, binds to the transactivation domain of the viral nuclear transcriptional activator, Tat, thereby increasing Tat's affinity for the transactivating response RNA element (TAR RNA). Serves as an essential cofactor for Tat, by promoting RNA Pol II activation, allowing transcription of viral genes.SUBUNIT Cyclin-T1 is the predominant cyclin that associates with CDK9 to form a heterodimer called P-TEFb (PubMed:30134174, PubMed:35393539, PubMed:9499409). P-TEFb forms a complex with AFF4/AF5Q31 (PubMed:12065898). Component of a complex which is at least composed of HTATSF1/Tat-SF1, P-TEFb complex, RNA pol II, SUPT5H, and NCL/nucleolin (PubMed:10393184). Component of the 7SK snRNP complex at least composed of P-TEFb (composed of CDK9 and CCNT1/cyclin-T1), HEXIM1, HEXIM2, BCDIN3, SART3 proteins and 7SK and U6 snRNAs (PubMed:17643375). Interacts (via central region) with ZMYND8 (via N-terminus); the interaction is direct and the association appears to occur between homodimeric ZMYND8 and the activated form of the P-TEFb complex (PubMed:30134174). Interacts with BRD4, targets chromatin binding (PubMed:16109376, PubMed:16109377, PubMed:24360279). Interacts with JMJD6 (PubMed:24360279). Interacts with MDFIC (PubMed:12944466). Interacts with HSF1 (PubMed:27189267). Interacts with HTATSF1 (PubMed:10913173). Interacts with TBX21 (PubMed:27292648).SUBUNIT (Microbial infection) Interacts with the transactivation region of HIV-1, HIV-2 and SIV Tat.SUBUNIT (Microbial infection) Interacts with human herpes virus 1 (HHV-1) transcriptional regulator ICP22.INTERACTION Ubiquitously expressed.DOMAIN The histidine-rich domain (HRD) region is intrinsically disordered and promotes the formation of phase-separated liquid droplets that enhance binding of the P-TEFb complex to the CTD (C-terminal domain) of the large subunit of RNA polymerase II (RNA Pol II).PTM ADP-ribosylation on serine residues by PARP1 in response to DNA damage disrupts the phase separation activity of CCNT1, thereby preventing activation of CDK9.SIMILARITY Belongs to the cyclin family. Cyclin C subfamily.
descriptionrecommendedName: fullName evidence="24"Cyclin-T1 shortName evidence="24"CycT1 shortName: Cyclin-T
geneNameCCNT1
identifierO60563
isSequenceChangedFALSE
keyword3D-structure
Acetylation
ADP-ribosylation
Alternative splicing
Cell cycle
Cell division
Coiled coil
Cyclin
Direct protein sequencing
Host-virus interaction
Isopeptide bond
Nucleus
Phosphoprotein
Proteomics identification
Reference proteome
Transcription
Transcription regulation
Ubl conjugation
modified[InstanceEdit:9836292] Weiser, Joel, 2023-05-25
[InstanceEdit:9852000] Weiser, Joel, 2023-11-03
[InstanceEdit:9917590] Weiser, Joel, 2024-08-09
[InstanceEdit:9926675] Weiser, Joel, 2024-11-03
nameCCNT1
referenceDatabase[ReferenceDatabase:2] UniProt
referenceGene[ReferenceDNASequence:8992936] ENSEMBL:ENSG00000129315 CCNT1 [Homo sapiens]
secondaryIdentifierCCNT1_HUMAN
A9XU13
E7EX76
O60581
sequenceLength726
species[Species:48887] Homo sapiens
(isoformParent)[ReferenceIsoform:8965092] UniProt:O60563-1 CCNT1 [Homo sapiens]
[ReferenceIsoform:8965093] UniProt:O60563-2 CCNT1 [Homo sapiens]
(referenceEntity)[EntityWithAccessionedSequence:51611] CCNT1 [nucleoplasm] [Homo sapiens]
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No pathways have been reviewed or authored by UniProt:O60563 CCNT1 (69859)