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Details on Person UniProt:O95292 VAPB

Class:IdReferenceGeneProduct:67106
_chainChangeLoginitiator methionine:1 added on Fri February 6 2015;chain:2-243 added on Fri February 6 2015;initiator methionine:1 for 67106 removed on Fri Nov 03 2023;initiator methionine: for 67106 added on Fri Nov 03 2023;initiator methionine: for 67106 removed on Fri Aug 15 2025;initiator methionine:1 for 67106 added on Fri Aug 15 2025
_displayNameUniProt:O95292 VAPB
_timestamp2025-08-15 21:50:16
chaininitiator methionine:1
chain:2-243
checksum22AEEF9EC7FC0B3F
commentFUNCTION Endoplasmic reticulum (ER)-anchored protein that mediates the formation of contact sites between the ER and endosomes via interaction with FFAT motif-containing proteins such as STARD3 or WDR44 (PubMed:32344433, PubMed:33124732). Interacts with STARD3 in a FFAT motif phosphorylation dependent manner (PubMed:33124732). Via interaction with WDR44 participates in neosynthesized protein export (PubMed:32344433). Participates in the endoplasmic reticulum unfolded protein response (UPR) by inducing ERN1/IRE1 activity (PubMed:16891305, PubMed:20940299). Involved in cellular calcium homeostasis regulation (PubMed:22131369).SUBUNIT Homodimer, and heterodimer with VAPA (PubMed:20940299). Interacts with VAMP1 and VAMP2 (PubMed:20940299, PubMed:9920726). Interacts (via MSP domain) with ZFYVE27 (PubMed:19289470, PubMed:21976701). Interacts with RMDN3 (PubMed:22131369). Interacts with KIF5A in a ZFYVE27-dependent manner (PubMed:21976701). Interacts (via MSP domain) with STARD3 (via phospho-FFAT motif) (PubMed:24105263, PubMed:33124732). Interacts with STARD3NL (via FFAT motif) (PubMed:24105263). Interacts with CERT1 (PubMed:16895911). Interacts with PLEKHA3 and SACM1L to form a ternary complex (PubMed:30659099). Interacts with VPS13A (via FFAT motif) (PubMed:30741634). Interacts with RB1CC1 (via phosphorylated FFAT motif), MIGA2 (via phosphorylated FFAT motif), RMDN3 (via phosphorylated FFAT motif), OSBPL1A (via FFAT motif), KCNB1 (via phosphorylated FFAT motif) and KCNB2 (via phosphorylated FFAT motif) (PubMed:33124732). Interacts (via MSP domain) with WDR44 (via FFAT motif); the interactions connect the endoplasmic reticulum (ER) with the endosomal tubule (PubMed:32344433).SUBUNIT (Microbial infection) Interacts (via MSP domain) with hepatitis C virus (HCV) non-structural protein 5A (via disordered domain D3) (PubMed:16227268, PubMed:22720086). Interacts with HCV RNA-directed RNA polymerase (PubMed:16227268).INTERACTION Present in mitochondria-associated membranes that are endoplasmic reticulum membrane regions closely apposed to the outer mitochondrial membrane.ALTERNATIVE PRODUCTS Ubiquitous. Isoform 1 predominates.DOMAIN The MSP domain binds the FFAT motif of many proteins.DISEASE The disease is caused by variants affecting the gene represented in this entry.DISEASE The disease is caused by variants affecting the gene represented in this entry.SIMILARITY Belongs to the VAMP-associated protein (VAP) (TC 9.B.17) family.ONLINE INFORMATION ALS genetic mutations dbONLINE INFORMATION Leiden Open Variation Database (LOVD)
descriptionrecommendedName: fullName evidence="22"Vesicle-associated membrane protein-associated protein B/C shortName: VAMP-B/VAMP-C shortName: VAMP-associated protein B/C shortName: VAP-B/VAP-C
geneNameVAPB
UNQ484/PRO983
identifierO95292
isSequenceChangedFALSE
keyword3D-structure
Acetylation
Alternative splicing
Amyotrophic lateral sclerosis
Coiled coil
Direct protein sequencing
Disease variant
Endoplasmic reticulum
Host-virus interaction
Isopeptide bond
Membrane
Neurodegeneration
Phosphoprotein
Proteomics identification
Reference proteome
Transmembrane
Transmembrane helix
Ubl conjugation
Unfolded protein response
modified[InstanceEdit:9836292] Weiser, Joel, 2023-05-25
[InstanceEdit:9852000] Weiser, Joel, 2023-11-03
[InstanceEdit:9862192] Weiser, Joel, 2024-02-26
[InstanceEdit:9917590] Weiser, Joel, 2024-08-09
[InstanceEdit:9926675] Weiser, Joel, 2024-11-03
[InstanceEdit:9963647] Weiser, Joel, 2025-08-15
nameVAPB
referenceDatabase[ReferenceDatabase:2] UniProt
referenceGene[ReferenceDNASequence:8994595] ENSEMBL:ENSG00000124164 VAPB [Homo sapiens]
secondaryIdentifierVAPB_HUMAN
A2A2F2
O95293
Q9P0H0
sequenceLength243
species[Species:48887] Homo sapiens
(isoformParent)[ReferenceIsoform:155626] UniProt:O95292-2 VAPB [Homo sapiens]
[ReferenceIsoform:402127] UniProt:O95292-1 VAPB [Homo sapiens]
(referenceEntity)[EntityWithAccessionedSequence:429718] VAPB [endoplasmic reticulum membrane] [Homo sapiens]
[EntityWithAccessionedSequence:9715115] VAPB [Golgi membrane] [Homo sapiens]
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No pathways have been reviewed or authored by UniProt:O95292 VAPB (67106)